All animal population growth rates are exponential (unless something catastrophic happens) so the population grows faster as it increases. For a general rule of thumb, it's common to see a wolf population grow by 300% in 5 to 10 years when you start with 110 breeding pairs. I ran in to one study that states that it's common to underestimate wolf population growth rates (depending upon how you monitor the population they say that underestimating by as much as 27% is not unusual). You can't help but wonder how much money was spent on that study - all they had to do was ask any rancher in the Idaho, Montana, Wyoming areas, they would have been able to tell the "experts" that they were underestimating the population growth rate several years ago.